- Born in 1929.
- Studied biological sciences in Freiburg and München.
- 1959
- PhD
- 1959 - 1962
- research grant to study rodent ethology.
- 1963 - 1967
- mammal section of the IRSAC Research Institute at Lwiro-Bukavu, East Congo: working on ecology and taxonomy of small mammals occurring in savannas and rain forests of Central Africa.
- 1967 - 1969
- as a curator of the mammal collection at the Museum Alexander Koenig at Bonn.
- Between 1969 and 1994
- head of mammal section at the SMNS. Continues to work on several research projects, i. e taxonomy and ecology of African rodents as well as mammals of Baden - Württemberg. About one hundred publications.
- Since 1994
- scientific volunteer at the Museum.